1QIMR Berghofer Medical Research Institute, Brisbane, QLD, Australia2National Cancer Institute, NIH, Bethesda, MD, USA3Weizmann Institute of Science, Rehovot, Israel4The University of Texas MD Anderson Cancer Center, Houston, TX, USA5School of Medicine, University of Colorado, Aurora, CO, USA6Melanoma Institute of Australia, Sydney, NSW, Australia7Tissue Pathology and Diagnostic Oncology, Royal Prince Alfred Hospital, Camperdown, NSW, Australia8Discipline of Pathology, The University of Sydney, Sydney, NSW, Australia9University of Sydney at Westmead Millenium Institute, Westmead, Sydney, NSW, Australia10Departments of Melanoma and Surgical Oncology, and Royal Prince Alfred Hospital, Sydney, NSW, Australia11Discipline of Surgery, Sydney Medical School, The University of Sydney, Sydney, NSW, Australia